@article {13116,
	title = {Description of the Larva of Quaestus (Quaesticulus) Pachecoi (Bolivar, 1915) (Coleoptera: Leiodidae: Cholevinae)},
	journal = {Annales Zoologici},
	volume = {61},
	year = {2011},
	pages = {691-708},
	abstract = {Two larval instars of Quaestus (Quaesticulus) pachecoi (Bolivar, 1915) are described and illustrated for the first time. Chaetotaxy, morphology and measurements indicate that the Q. (Q.) pachecoi has a classic type of development with two instars only and larvae belong to a classic type according to the Deleurance-Gla{\c c}on classification.},
	keywords = {cave beetles, chaetotaxy, Cholevinae, classic type of development, classic type of larvae, glands, Iberian Peninsula, larva, Leiodidae, Leptodirini, morphology, Quaestus (Quaesticulus) pachecoi},
	author = {Adamczyk, B and Kilian, A and Salgado, JM}
}
@article {13065,
	title = {A chromosomal investigation of some European Leiodidae (Coleoptera), with particular focus on Spanish subterranean Leptodirini},
	journal = {CompCytogen},
	volume = {6},
	year = {2012},
	pages = {127{\textendash}139 },
	abstract = {Karyotypes are shown for Leiodes calcarata (Erichson, 1845), Catops coracinus Kellner, 1846, Cantabrogeus
luquei (Salgado 1993), Espanoliella luquei Salgado \& Fresneda, 2005, Fresnedaella lucius Salgado, Labrada
\& Luque 2011, Notidocharis uhagoni (Sharp, 1872), Quaestus (Quaesticulus) pasensis Salgado, Labrada \&
Luque, 2010, all of which are shown to have a diploid number of 20 autosomes plus Xy (♂) or XX (♀)
sex chromosomes, as well as an as yet undescribed triploid species of the genus Cantabrogeus Salgado,
2000. These results are contrasted with published information, all on Leptodirini, which lists 10 species
as having diploid numbers of 22 + Xy or XX. It is shown that the higher chromosome number (n = 11 +
X or y) previously reported refers exclusively to the more derived Leptodirini ({\textquotedblleft}infraflagellates{\textquotedblright}) whereas
the lower number (n = 10 + X or y) refers to the less derived surface-dwelling forms and the less derived
Leptodirini ({\textquotedblleft}supraflagellates{\textquotedblright}).},
	keywords = {1796, 1798, 1861, 1956, 2011, Cantabria, Cantabrogeus Salgado, Catops Paykull, Chromosomes, Jeannel, Labrada \& Luque, Leiodes Latreille, Leiodidae, Leptodirini, Notidocharis, Quaestus Schaufuss, triploid},
	author = {Angus, R.B. and Edwards, D.B. and C. G. Luque and Labrada, L}
}
@article {10155,
	title = {The phylogenetic and geographic limits of Leptodirini (Insecta: Coleoptera: Leiodidae: Cholevinae), with a description of Sciaphyes shestakovi sp. n. from the Russian Far East},
	journal = {Arthropod Systematics \& Phylogeny},
	volume = {69},
	year = {2011},
	pages = {99 {\textendash}123},
	abstract = {The tribe Leptodirini of the beetle family Leiodidae is one of the most diverse radiations of cave animals, with a distribution 
centred north of the Mediterranean basin from the Iberian Peninsula to Iran. Six genera outside this core area, most notably 
Platycholeus Horn, 1880 in the western United States and others in East Asia, have been assumed to be related to Leptodirini. We studied recently collected specimens of three of these extraterritorial genera, namely Platycholeus, Fusi Perkovsky, 
1989 and Sciaphyes Jeannel, 1910, and establish their phylogenetic relationships by analysing a combination of ca. 5 Kb of 
mitochondrial and nuclear DNA sequences with Bayesian Probability and Maximum Likelihood methods. Our results corroborate the previously proposed hypothesis that Platycholeus is the sister group of the remaining Leptodirini, with an estimated age of vicariant separation compatible with the breaking of the Thulean bridge between the Nearctic and the Western 
Palaearctic in the Early Eocene. We refute close relationship of either Fusi or Sciaphyes to Leptodirini, with the former genus appearing more closely related to Cholevini, and the latter to Anemadini and warranting a separate tribe, Sciaphyini, in 
agreement with recent treatments. This phylogenetic position of Sciaphyes is in agreement with a parsimony analysis of 
28 morphological characters of a representative sample of Cholevinae subgroups. We describe one of the studied Sciaphyes
species as S. shestakovi sp.n. and indicate its remarkable morphological differences from its congeners. We hypothesise that 
the remaining three extraterritorial monotypic genera of {\textquotedblleft}Leptodirini{\textquotedblright}, namely Proleptodirina Perkovsky, 1998, Sinobathys-
cia Perreau, 1999 and Coreobathyscia Szymczakowski, 1975 are unlikely to be closely related to the tribe, which probably 
has its easternmost geographical limits at Zagros and Alborz (= Elburz) mountains in Iran.
},
	keywords = {Fusi, Leptodirini, phylogeny, Platycholeus, Russia, Sciaphyes shestakovi sp. n., Siberia, subterranean environment},
	author = {Fresneda, J and Grebennikov, V. V. and Ribera, I}
}
