@inbook {13253,
	title = {G{\'e}nero Ptomaphagus Hellwig, 1795},
	booktitle = {Fauna Iberica Vol. 31 Coleoptera Cholevidae},
	year = {2008},
	pages = {696-708},
	author = {Salgado Costas, JM and Blas, M and Fresneda, J}
}
@article {10852,
	title = {Speonomus kryophilos n. sp. (Coleoptera, Cholevidae) nuevo Bathysciinae del Pirineo Catal{\'a}n},
	journal = {Ilerda Ci{\`e}ncies},
	volume = {49},
	year = {1991},
	pages = {259-264.},
	author = {Fresneda, J and Hernando, C}
}
@article {10758,
	title = {Baronniesia delioti gen. n. sp. n., a new subterranean Leptodirini from the French Pyrenees (Coleoptera: Leiodidae: Cholevinae)},
	journal = {Zootaxa},
	volume = {1},
	year = {1993},
	pages = {2009},
	author = {Fresneda, J and Bourdeau, C and Faille, A}
}
@article {10621,
	title = {Revisi{\'o}n de los Speonomus Jeannel, 1908 de los grupos bolivari y brieti (Coleoptera: Cholevidae, Leptodirinae)},
	journal = {Zapateri, Revista aragonesa de Entomolog{\'\i}a},
	volume = {8},
	year = {1999},
	pages = {121-156.},
	author = {Fresneda, J and Hernando, C and Lagar, A}
}
@article {10547,
	title = {Stygiophyes hansferyi n. sp. del Pirineo de Huesca y L{\'e}rida (Espa{\~n}a) (Col., Cholevidae, Leptodirinae)},
	journal = {MBsp},
	volume = {28},
	year = {2002},
	pages = {71-77.},
	author = {Fresneda, J and Escol{\`a}, O}
}
@article {10491,
	title = {The genus Espanoliella Gu{\'e}orguiev, 1976. E. luquei, sp. nov. (Coleoptera: Leiodidae: Leptodirinae)},
	journal = {STbl},
	volume = {3},
	year = {2005},
	pages = {87-96.},
	author = {Salgado, JM and Fresneda, J}
}
@article {10487,
	title = {The genus Bathysciola Jeannel, 1910 in the iberian peninsula and Pyrenees, taxonomic revision of sections IV, VI and VII (Jeannel, 1924) (Coleoptera, Cholevidae, Leptodirinae)},
	journal = {Graellsia},
	volume = {62},
	number = {1},
	year = {2006},
	pages = {25-74.},
	author = {Fresneda, J and Salgado, JM}
}
@article {10459,
	title = {Datos de distribuci{\'o}n de Speonemadus bolivari (Jeannel, 1922) (Coleoptera, Leiodidae, Cholevinae, Anemadini)},
	journal = {Monograf{\'\i}as Biospeleol{\'o}gicas},
	volume = {3},
	year = {2008},
	pages = {13-15.},
	author = {Fresneda, J}
}
@article {10451,
	title = {Eskualdunella delespierrei Coiffait, 1950: la cl{\'e} inattendue d{\textquoteright}une {\'e}nigme biog{\'e}ographique (Coleoptera, Leiodidae)},
	journal = {BSef},
	volume = {114},
	number = {4},
	year = {2009},
	pages = {469-473.},
	author = {Faille, A and Bourdeau, C and Fresneda, J}
}
@article {10439,
	title = {El g{\'e}nero Trapezodirus Jeannel, consideraciones biogeogr{\`a}ficas y presencia en el medio hipogeo del macizo de Cotiella (Pirineos, Huesca, Espa{\~n}a) (Coleoptera, Leiodidae, Cholevinae, Leptodirini)},
	journal = {Cotiella},
	volume = {1},
	year = {2009},
	pages = {24-40.},
	author = {Fresneda, J and Bourdeau, C and Faille, A}
}
@article {10440,
	title = {Baronniesia delioti gen. n. sp. n., a new Leptodirini from the French Pyrenees (Coleoptera: Leiodidae: Cholevinae)},
	journal = {Zootaxa},
	volume = {1993},
	year = {2009},
	pages = {1-16.},
	author = {Fresneda, J and Bourdeau, C and Faille, A}
}
@article {10428,
	title = {Ancient origin of a Western  Mediterranean radiation of subterranean beetles},
	journal = {BMC Evolutionary Biology},
	volume = {10},
	number = {29},
	year = {2010},
	pages = {1-14arab.},
	chapter = {29},
	abstract = {Background: Cave organisms have been used as models for evolution and biogeography, as their reduced above- ground dispersal produces phylogenetic patterns of area distribution that largely match the geological history of mountain ranges and cave habitats. Most current hypotheses assume that subterranean lineages arose recently from surface dwelling, dispersive close relatives, but for terrestrial organisms there is scant phylogenetic evidence to support this view. We study here with molecular methods the evolutionary history of a highly diverse assemblage of subterranean beetles in the tribe Leptodirini (Coleoptera, Leiodidae, Cholevinae) in the mountain systems of the Western Mediterranean.
Results: Ca. 3.5 KB of sequence information from five mitochondrial and two nuclear gene fragments was obtained for 57 species of Leptodirini and eight outgroups. Phylogenetic analysis was robust to changes in alignment and reconstruction method and revealed strongly supported clades, each of them restricted to a major mountain system in the Iberian peninsula. A molecular clock calibration of the tree using the separation of the Sardinian microplate (at 33 MY) established a rate of 2.0\% divergence per MY for five mitochondrial genes (4\% for cox1 alone) and dated the nodes separating the main subterranean lineages before the Early Oligocene. The colonisation of the Pyrenean chain, by a lineage not closely related to those found elsewhere in the Iberian peninsula, began soon after the subterranean habitat became available in the Early Oligocene, and progressed from the periphery to the centre.
Conclusions: Our results suggest that by the Early-Mid Oligocene the main lineages of Western Mediterranean Leptodirini had developed all modifications to the subterranean life and were already present in the main geographical areas in which they are found today. The origin of the currently recognised genera can be dated to the Late Oligocene-Miocene, and their diversification can thus be traced to Miocene ancestors fully adapted to subterranean life, with no evidence of extinct epigean, less modified lineages. The close correspondence of organismal evolution and geological record confirms them as an important study system for historical biogeography and molecular evolution.},
	author = {Ribera, I and Fresneda, J and Bucur, R and Izquierdo, A. and Vogler, AP and Salgado, JM and Cieslak, A.}
}
@article {10424,
	title = {Un nuevo troglobio de la region Cantabrica: Quaestus (Speogeus) jubilationis n. sp. (Coleoptera: Leiodidae: Cholevinae: Leptodirini)},
	journal = {HTpt},
	volume = {10},
	number = {2},
	year = {2010},
	pages = {99-106.},
	abstract = {A new cave-dwelling species from the Cantabrian region: Quaestus (Speogeus) jubilationis n. sp. (Coleoptera: Leiodidae: Cholevinae: Leptodirini)
A new species belonging to the subgenus Speogeus Salgado, 1985, Queastus (Speogeus) jubilationis n. sp. is described. The taxonomic position is discussed, based on morphological characters, the aedaegus and the spermathecal complex. A new identification key to the species of the subgenus is proposed.},
	keywords = {Cantabrian region, Coleoptera, distribution, Leiodidae, Queastus (Speogeus) jubilationis n. sp., Spain, taxonomy},
	author = {Salgado, JM and Fresneda, J}
}
@article {10419,
	title = {Le genre Paratroglophyes Four{\`e}s, 1954: description d{\textquoteright}une nouvelle esp{\`e}ce du massif du mont Valier, Pyr{\'e}n{\'e}es centrales, France (Coleoptera, Leiodidae, Cholevinae, Leptodirini)},
	journal = {BSef},
	volume = {116},
	number = {3},
	year = {2011},
	pages = {355-363.},
	author = {Fresneda, J and Bourdeau, C and Faille, A}
}
@article {10219,
	title = {Descripci{\'o}n del primer Catops Paykull, 1798 end{\'e}mico de la Pen{\'\i}nsula Ib{\'e}rica: Catops punctatulus n. sp. (Coleoptera, Leiodidae, Cholevinae)},
	journal = {Nouvelle Revue d{\textquoteright}Entomologie},
	volume = {23(1)},
	year = {2006},
	pages = {45-54},
	author = {Fresneda, J and Lencina, JL and Salgado, JM}
}
@article {10190,
	title = {The genus Bathysciola Jeannel, 1910 in the Iberian Peninsula and Pyrenees, taxonomic revision of sections IV, VI and VII (Jeannel, 1924) (coleoptera, cholevidae, leptodirinae)},
	journal = {Graellsia: revista de zoolog{\'\i}a},
	volume = {Vol. 62, N{\textordmasculine}. 1 },
	year = {2006},
	pages = {25-74},
	author = {Salgado, JM and Fresneda, J}
}
@article {10185,
	title = {Descripci{\'o}n de Bathysciola liqueana sp. n. de los Pirineos centrales (Francia). Designaci{\'o}n de lectotipos y datos de distribuci{\'o}n de las especies del grupo de B. meridionalis (Jacquelin du Val, 1854) (Insecta, Coleoptera, Leiodidae, Cholevinae, Leptodirin},
	journal = {Animal Biodiversity and Conservation},
	volume = {Vol.: 33 N{\'u}m.: 2},
	year = {2010},
	pages = {131-142},
	author = {Fresneda, J and Bourdeau, C and Faille, A}
}
@article {10170,
	title = {Une nouvelle esp{\`e}ce pyr{\'e}n{\'e}enne et une nouvelle synonymie dans le genre Bathysciola Jeannel, 1910 (Coleoptera, Leiodidae, Cholevinae, Leptodirini) },
	journal = {Bulletin de la Soci{\'e}t{\'e} entomologique de France},
	volume = {115 (4), 2010 : 427-433.},
	year = {2010},
	author = {Fresneda, J and Perreau, M. and Vandenbergh, C}
}
@article {10155,
	title = {The phylogenetic and geographic limits of Leptodirini (Insecta: Coleoptera: Leiodidae: Cholevinae), with a description of Sciaphyes shestakovi sp. n. from the Russian Far East},
	journal = {Arthropod Systematics \& Phylogeny},
	volume = {69},
	year = {2011},
	pages = {99 {\textendash}123},
	abstract = {The tribe Leptodirini of the beetle family Leiodidae is one of the most diverse radiations of cave animals, with a distribution 
centred north of the Mediterranean basin from the Iberian Peninsula to Iran. Six genera outside this core area, most notably 
Platycholeus Horn, 1880 in the western United States and others in East Asia, have been assumed to be related to Leptodirini. We studied recently collected specimens of three of these extraterritorial genera, namely Platycholeus, Fusi Perkovsky, 
1989 and Sciaphyes Jeannel, 1910, and establish their phylogenetic relationships by analysing a combination of ca. 5 Kb of 
mitochondrial and nuclear DNA sequences with Bayesian Probability and Maximum Likelihood methods. Our results corroborate the previously proposed hypothesis that Platycholeus is the sister group of the remaining Leptodirini, with an estimated age of vicariant separation compatible with the breaking of the Thulean bridge between the Nearctic and the Western 
Palaearctic in the Early Eocene. We refute close relationship of either Fusi or Sciaphyes to Leptodirini, with the former genus appearing more closely related to Cholevini, and the latter to Anemadini and warranting a separate tribe, Sciaphyini, in 
agreement with recent treatments. This phylogenetic position of Sciaphyes is in agreement with a parsimony analysis of 
28 morphological characters of a representative sample of Cholevinae subgroups. We describe one of the studied Sciaphyes
species as S. shestakovi sp.n. and indicate its remarkable morphological differences from its congeners. We hypothesise that 
the remaining three extraterritorial monotypic genera of {\textquotedblleft}Leptodirini{\textquotedblright}, namely Proleptodirina Perkovsky, 1998, Sinobathys-
cia Perreau, 1999 and Coreobathyscia Szymczakowski, 1975 are unlikely to be closely related to the tribe, which probably 
has its easternmost geographical limits at Zagros and Alborz (= Elburz) mountains in Iran.
},
	keywords = {Fusi, Leptodirini, phylogeny, Platycholeus, Russia, Sciaphyes shestakovi sp. n., Siberia, subterranean environment},
	author = {Fresneda, J and Grebennikov, V. V. and Ribera, I}
}
